Dendroculus Guide: Find All Locations in Genshin Impact Sumeru

There are 271 Dendroculus to discover across Sumeru in Genshin Impact. This comprehensive Dendrooculus Guide provides interactive maps and detailed instructions for locating every Dendroculus in Sumeru’s Rainforest, Desert, Hadramaveth, and Girdle of the Sands regions. Collect them all to upgrade the Statues of the Seven and earn valuable rewards!

Elemental Oculus Items
Anemoculus
Dendroculus

Interactive Dendroculus Maps

This guide is structured to help you efficiently find all Dendroculus. Use the maps below to get a visual overview of each region, then follow the detailed descriptions for each Dendroculus location.

Dendroculus Locations in Each Region

Rainforest Dendroculus	Great Red Sand Dendroculus
Girdle of the Sands Dendroculus
Tip: Utilize your browser’s page search function (Ctrl+F or Cmd+F) to quickly find a specific Dendroculus number from the maps!

As of Version 3.6, there are a total of 271 Dendroculi scattered throughout Sumeru.

Explore More Dendroculus Guides
Rainforest Dendroculus
Hadramaveth Dendroculus

Rainforest Dendroculus Locations {#rainforest-dendroculus-locations}

No.	Image	How to Get
1		During the “The Final Chapter” quest of the Agnihotra Sutra quest chain, reach the final cave and proceed to the deepest point. You’ll see a rock below. Use any Dendro attack to hit the rock. It will rise, making the Dendroculus accessible.
2		In Matiyama Forest, find it inside a giant mushroom/tree. Climb to the top to collect.
3		Located in the northernmost part of Sumeru, on top of a giant mushroom. Obtain Dendrograna and shoot at the three floating targets to remove the barrier and claim the Dendroculus.
4		In Mawtiyima Forest, activate the mushroom on the ground near Anemo Slimes to create a wind current. Ride the current to reach the Dendroculus high in the sky.
5		On a mushroom inside a cave beneath the waypoint in Mawtiyima Forest. The cave entrance is southwest of the waypoint.
6		In Matiyama Forest, on top of another giant mushroom/tree. Climb the mushroom to get it.
7		South of Mawtiyima Forest, on top of a giant boulder. Climb the boulder to collect the Dendroculus.
8		At the entrance of the cave leading to the Electro Regisvine boss.
9		On top of an umbrella on the hill where the Electro Regisvine’s cave is located. Use Dendro on the nearby Clusterleaves of Cultivation to create a Four-Leaf Sigil and reach the Dendroculus.
10		North of Gandha Hill, between overhanging roots. Use the Four-Leaf Sigil to access it.
11		In Lokapala Jungle, south of Mawtiyima Forest, on top of a large mushroom with three Anemo Slimes. Use Four-Leaf Sigils to ascend the mushrooms.
12		At the top of the Palace of Alcazarzaray. Climb the palace or glide from the cliffs near the northeast teleport waypoint.
13		By the waterfalls west of Gandha Hill. Jump from the top of the waterfalls or use the Bouncy Mushrooms around to reach the Dendroculus.
14		Under a ledge in Chatrakam Cave. Jump and glide down to get it.
15		In Chatrakam Cave. Only accessible during the “Risen Moon Chapter” quest of Agnihotra Sutra. Requires the upgraded Kusava gadget.
16		Inside a cave during the Agnihotra Sutra quest. Hit all floating rings with Dendrograna to break the barrier.
17		Accessible from the Agnihotra Sutra, “Risen Moon Chapter” quest. Progress through the quest to find it.
18		Apply Electro to the Bouncy Mushroom to spawn Four-Leaf Sigils. Use the mushroom and sigils to reach the Dendroculus.
19		Clear the Withering zone first, then apply Electro to the Bouncy Mushroom to reach the Dendroculus.
20		On top of a tree. Use the Four-Leaf Sigils to reach it quickly.
21		Inside a cave southeast of Vanarana. Unflood the area by destroying two Withering Branches, then activate the Dendro Monument.
22		In an underground cave west of Sumeru City. Find the tunnel east of the waypoint and follow the path.
23		In a cave with Hydro Slimes south of Vanarana, near a Statue of The Seven.
24		By the road south of Vanarana, inside a hut.
25		Southeast of a Statue of Seven in Vanarana, in a clearing above a pond. Use the Four-Leaf Sigil or glide from a higher area.
26		Just before the waterfall in Vanarana, in a small hut.
27		Inside the hollow space of a big tree trunk in Vanarana. Use Four-Leaf Sigils for easy access.
28		In a cave where a small farm is located in Vanarana. Enter the cave and go right at the fork.
29		By the Jadeplume Terrorshroom boss location, underneath a stone arc leading to Shroom-Kin enemies.
30		South of the Jadeplume Terrorshroom boss location. Floating above the hole leading down to the boss.
31		Under a rock. Clear the Withering zone to unearth it.
32		Above a waterfall. Use the Bouncy Mushroom on one side to reach it.
33		North of Yasna Monument, high in the sky. Use Electro on the Bouncy Mushroom on the large boulder with a Hilichurl Camp.
34		Clear the Withering zone to revitalize the Bouncy Mushroom. Use Electro for a higher bounce!
35		Inside the cave where you need to lower the water level in Yasna Monument area.
36		Underground, during the Aranayaka Part III quest. Progress through the quest to find it.
37		North of Pardis Dhyai, underneath a rock. Appears after clearing the Withering zone.
38		Solve puzzles inside the cave to reach a flooded underground area. The Dendroculus is on a trunk. Use songs from the Vintage Lyre.
39		South of Yazadaha Pool, underneath a stone bridge with large roots. Jump from the bridge to get it.
40		Left from Yazadaha Pool, high up. Use the Bouncy Mushroom below it and Electro for extra height.
41		Near Yazadaha Pool, high up. Clear the Withering zone to revitalize the Bouncy Mushroom. Use Electro on it.
42		West of Sumeru city, at the end of big tree roots. Use the Bouncy Mushroom below; use Electro to jump higher.
43		At the tip of a branch of the giant tree in Sumeru City. Climb Sanctuary of Surastana and onto the tree trunk.
44		Under the large wood bridge southwest of Chinvat Ravine. Jump from the bridge or use the Bouncy Mushroom.
45		Past the large wood bridge, underneath a dendro barrel. Explode the barrel with Pyro.
46		West of Gandharva Ville, inside a tree trunk house.
47		On top of the giant tree house at Gandharva Ville. Follow the path up and climb the tree trunk.
48		On a rocky arch by the road at the northeast entrance of Gandharva Ville. Use Venti/Kazuha or Geo Constructs.
49		Complete the Agnihotra Sutra quest for the “Kusava” gadget. Fix the broken monument to clear the roots.
50		On top of an entangled boulder. Climb the boulder to reach it.
51		Under a rock after clearing the Withering zone.
52		Inside a cave unlocked during the Vimana Agama: Rojinyan’s Chapter quest.
53		At the edge of a cliff south of Vimara Village, with a Hilichurl Shooter below. Glide down carefully.
54		Inside a ruined hut at the top of the hill near Vimara Village.
55		On a large gear under a rock arch. Climb the rocks to reach the gear.
56		Underwater in an underground cave. Plunge attack from the branch above to grab it without draining water.
57		Close to the Liyue-Sumeru border, southwest of the teleport waypoint by Tiangong Gorge.
58		Summon Dendrograna and shoot the floating target with a Charged Bow Attack. Repeat for the three targets it summons to lower the barrier.
59		On a mound beside a tree near the Liyue border.
60		Play Rhythm of the Beastly Trail on the flower with the Vintage Lyre, or simply touch/stand on the rock.
61		On a tree east of Devantaka Mountain. Climb the tree to collect it.
62		On top of a gear in the middle of Devantaka Mountain. Glide from the Statue of the Seven (Ardravi Valley, Sumeru).
63		Inside one of the fingers of the massive corroded robot in Devantaka Mountain.
64		East of Devantaka Mountains, inside a hollowed tree trunk.
65		South of Devantaka Mountain, under the arm of the massive corroded robot.
66		Inside the giant robot’s eye in Devantaka Mountain. Run along its arm or glide from the south waypoint.
67		Southeastern mountain range of Sumeru, east of Devantaka Mountains, inside a Dendro dome. Pick up flowers to spawn Whopperflowers and lower the barrier.
68		Southeast area of Devantaka Mountain. Hidden in tree leaves, guarded by an enemy. Scale the mountain.
69		Underground south of Devantaka Mountain. Enter from the north, head south to find giant metal parts. Use Electro on the Bouncy Mushroom to reach it.
70		East of Port Ormos, inside a Dendro dome. Play Rhythm of the Beastly Trail to unlock it.
71		At the mouth of a river north of Port Ormos. On a net beside an inactive Ruin Guard.
72		In the middle of the lake north of Port Ormos. Use a waverider from the nearby waypoint.
73		On a large gear north of Port Ormos. Climb the structure to reach the gear.
74		Inside a hollow trunk west of Port Ormos. Enter the trunk to find it.
75		On the roof of a hut northwest of Port Ormos. Climb the hut.
76		Inside a giant log southeast of Apam Woods. Enter the log.
77		Clear the Withering southwest of Vimara Village. Find the bouncy mushroom below the Dendroculus and energize it with Electro.
78		In an alcove at the base of the valley between Pardis Dhyai and Vimara Village. A Hilichurl is digging on top.
79		Beneath a large hollow tree trunk overlooking the river between Pardis Dhyai and Vimara Village. Climb the tree trunk.
80		Under the bridge south of Pardis Dhyai. Glide down to collect it.
81		Under a rock after clearing the Withering zone north of Apam Woods.
82		At the base of a tree at the entrance of a Withering zone in Apam Woods.
83		In the middle of a waterfall in Apam Woods. Climb to the top of the waterfall.
84		Under a rock after clearing a Withering zone east of Apam Woods.
85		Beneath a tree at the center of Apam Woods. Go up the slope on the west and glide.
86		At the top of a waterfall in Apam Woods. Use Clusterleaves of Cultivation for a Four-Leaf Sigil.
87		From the Tower of Abject Pride domain, glide east to a platform. Use Bouncy Mushroom and Four-Leaf Sigils to a higher platform. Use Dendro on Clusterleaves for more sigils.
88		Under a plant after clearing the Withering zone north of the Tower of Abject Pride domain.
89		After completing the four subquests of Varuna Gatha quest chain. South of Apam Woods on an island in the lake.
90		On a large tree southeast of Apam Woods. Use the Bouncy Mushroom near the waypoint.
91		Under a giant root north of The Coordinates of Sun and Rain domain. Climb the root and glide down.
92		Under a plant after clearing the Withering zone southwest of The Coordinates of Sun and Rain domain.
93		Under a wooden platform southeast of the Tower of Abject Pride domain.
94		In Grove of Dreams (underground), on the topmost branch of the large tree.
95		East of Ruins of Dahri, under a plant holding a rock. Clear the Withering zone.
96		Down the hole in Ruins of Dahri. Access via the “Irate Iron Chunk” quest of Varuna Gatha.
97		South of Ruins of Dahri, near the waypoint. Climb the overgrowth to the tip of a root.
98		Play Rhythm of the Gloomy Path. Enter the portal.
99		At the top of the giant tree in Ruins of Dahri. Climb the tree.
100		Play Rhythm of the Gloomy Path. Enter the portal.
101		Under a rock after clearing a Withering zone north of Caravan Ribat.
102		Do “The Rhythm that Nurtures the Sprout” quest. Use Vintage Lyre and shoot the floating target.
103		On top of a tower left of Yasna Monument. Glide from the closest waypoint.
104		In Yasna Monument. Revive Dendrograna Branch, hit floating Dendro rings with Healthy Dendrograna.
105		Above a stone statue on top of a hill near Yasna Monument.
106		Clear the Withering zone to unearth it from under a stone.
107		Clear the Withering zone to unearth it from under a stone.
108		North of Yasna Monument, on top of a large tree. Climb the tree.
109		Complete “For All Children Who Long for Life” to explore Old Vanarana safely. Apply Dendro to a rock to reveal it.
110		Complete “For All Children Who Long for Life” to explore Old Vanarana safely. On top of a spiral dead tree.

Great Red Sand Desert Dendroculus Locations {#great-red-sand-desert-dendroculus-locations}

No.	Image	How to Get
1		West of the Eye of the Sand, above the gate next to a teleport waypoint.
2		Above a remarkable chest. Accessible during the Dreams Beneath the Searing Sand World Quest.
3		In the same room as an Obscuring Cell and Primal Beam during the Dreams Beneath the Searing Sand World Quest. Behind a crumbled wall on the north side.
4		At the end of the invisible maze inside Khaj-Nisut. Only during Dreams Beneath the Searing Sand World Quest.
5		After revealing Khaj-Nisut during Dreams Beneath the Searing Sand World Quest. Glide from the southeast waypoint to the top of Khaj-Nisut.
6		Behind a statue in the invisible maze room in Khaj-Nisut. Only during Dreams Beneath the Searing Sand World Quest.
7		East of the Eye of the Sand, at the top of a rock formation. Climb the formation.
8		East of The Dune of Carouses, behind three wooden boxes below the cliff.
9		By Abdju Pit, west of a small camp. Found by following the Golden Slumber World Quest.
10		Near a common chest circled by Geo Fungi and a Geoshroom. Defeat the enemies.
11		At the hole leading underground, drop down the gale to grab it.
12		In front of a ruined tower in Aaru Village.
13		At the top of the building in the center of Aaru Village, next to the waypoint.
14		By a hole in Dar al-Shifa. Explore the area near the hole.
15		After gaining clearance through the Golden Slumber Questline, return to the ruins near Abdju Pit. Activate the plinth and follow the path to the room with Primal Construct: Reshaper. It’s along a path past this room.
16		In the ruins during the Lost in the Sands quest. Progress through the quest.
17		Deeper in the ruins near Abdju Pit, in the room where the elevator from Dendroculus #19 takes you.
18		Inside the Plint of the Secret Rites room during the Lost in the Sands quest. Entrance from the hidden waypoint near the Domain on the top left.
19		Following the path from Lost in the Sands quest, find a second plinth. Operate it after gaining clearance to call an elevator.
20		At the top of an Ajilenakh Nut tree. Use the Four-Leaf Sigil and glide down.
21		Destroy rocks beneath Dendroculus #14 for an underground cave. Follow the path, breaking rocks, to a Hilichurl camp. Destroy rocks behind the camp.
22		Just below a cliff east of the Garden of Endless Pillars domain. Glide down carefully.
23		Jump down the pit south of the waypoint and find an opening beneath the statue. Follow the path north into the tower. At the top, it’s behind the pedestal.
24		Inside The Dune of Carouses, change the Prism of Khaj-Nisut’s color to blue. This disables the invisible barrier to the left.
25		Ride the upward draft south of The Dune of Carouses and follow the wind hoops.
26		Southeast of The Dune of Carouses, below a ledge northwest of a Statue of the Seven.
27		Glide on the upward draft near the teleport waypoint above an invisible barrier.
28		In ruins northwest of The Mausoleum of King Deshret. Unlocked in The Secret of Al-Ahmar quest.
29		On a crumbled wall near an Exquisite Chest guarded by Primal Constructs. Defeat the enemies.
30		North of The Mausoleum of King Deshret. Unlocked in The Secret of Al-Ahmar World Quest.
31		Near Primal Torches during The Secret of Al-Ahmar World Quest. Complete the Primal Sandglass Puzzle for a Four-Leaf Sigil.
32		Gain clearance then fast travel to Khemenu Temple. Head west to the elevator room. In the north room, follow the path to a plinth. Operate it to open a door.
33		In buildings unlocked in An Introduction to Indoor Archaeology quest. Progress through the quest.
34		On the west end of Khemenu Temple’s invisible ceiling. Explore the upper areas.
35		Outside of The Mausoleum of King Deshret.
36		In front of a giant fan in the Place of Breath Puzzle of Dual Evidence World Quest.
37		In buildings unlocked in An Introduction to Indoor Archaeology quest.
38		In the middle of a broken bridge in the Place of Breath Puzzle. Dual Evidence World Quest, requires Lv. 2 clearance.
39		In Place of Breath Puzzle ruins. Move primal beams to activate Anemo monuments, summon an updraft.
40		In The Mausoleum of King Deshret. After completing The Secret of Al-Ahmar World Quest.
41		In The Mausoleum of King Deshret. Unlocked in The Secret of Al-Ahmar World Quest.
42		At the top of The Mausoleum of King Deshret. Climb to the very top.
43		In The Mausoleum of King Deshret. Unlocked in The Secret of Al-Ahmar World Quest.
44		Above statues in a cavern accessed via The Secret of Al-Ahmar World Quest. Exit from the southwest waypoint and use Four-Leaf Sigil.
45		After Lv.3 clearance, solve the Primal Sandglass Puzzle and light the Primal Torch to remove invisible walls.
46		Activate the giant fan after Lv.3 clearance and glide up.
47		On a tree in an underground Withering zone. Cavern accessed via The Secret of Al-Ahmar World Quest.
48		From the waypoint by Dendroculus #50, face northwest and find a tunnel above. Follow the path to a room with giant mushrooms. On the biggest mushroom.
49		Entrance to ruins in the southwest desert. Follow the path and solve puzzles, ascend via elevator.
50		By the entrance to ruins south of The Mausoleum of King Deshret. Visible from the nearby waypoint.
51		Gain clearance and enter The Dune of Magma. In a pit beneath, past a remarkable chest.
52		On scaffolding in the Place of Breath Puzzle. Interact with rocks to summon an updraft.
53		At the top of the pyramid in The Dune of Elusion. Use Four-Leaf Sigils on west and south walls.
54		Inside ruins south of The Dune of Elusion. West end of ruins, requires Lv.3 clearance.
55		South of The Dune of Elusion. Activate three Pyro monuments to lower the barrier.
56		East of The Dune of Elusion, on a tree surrounded by Pyro Fungi.
57		South of Sobek Oasis, floating by a cliff east of a teleport waypoint.
58		Southeast of Sobek Oasis, above a Fatui tent. Climb the rocks nearby.
59		North of the Valley of Dahri, above a pond in an underground cave north of a Statue of the Seven.
60		West of the Valley of Dahri, on a branch by a tower.
61		Inside an Eremite hideout. Follow the Female Eremite for the password, use it on the mechanism at the entrance. First room to the right.
62		By the Valley of Dahri, on a branch by a cliff. Use Four-Leaf Sigils from the teleport waypoint.
63		At the back of the Ruin Golem. Climb to the back of the golem.
64		Inside the Ruin Golem during Afratu’s Dilemma World Quest. On the wall next to a Suda’s Flow Seal.
65		Inside the Ruin Golem’s right arm, behind a giant gear. Accessible from the main floor.
66		After disabling the Ruin Golem’s auto-fire in Afratu’s Dilemma World Quest. Fire at boulders on the right to open a cavern.
67		In an underground cave with a Withering zone blocked by a boulder. Destroy boulder using the Ruin Golem in Afratu’s Dilemma World Quest.
68		Northeast of the Valley of Dahri. Use Four-Leaf Sigils above the teleport waypoint.
69		In ruins during the Lost in the Sands quest. Building entrance unlocked from inside during the quest.
70		Above the temple entrance northeast of Sobek Oasis, next to the teleport waypoint.

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Desert of Hadramaveth Dendroculus Locations {#desert-of-hadramaveth-dendroculus-locations}

No.	Image	How to Get
1		Behind a rock wall, near the Fatui Stronghold gate. Break the rock wall.
2		Atop a ruined pillar, north of the waypoint. Climb the pillar.
3		Floating in the air, near the right arm of the Colossal Ruin Guard. Glide from a higher point.
4		Near an Eremite camp, under the Colossal Ruin Guard. Check under the guard’s structure.
5		Above the Debris of Panjvahe waypoint. Use the Four-Leaf Sigil to reach it.
6		Near a ruin guard, north of the waypoint in Debris of Panjvahe. Defeat the guard or avoid it.
7		On top of a rock, near the fissure in Debris of Panjvahe. Climb the rocks near the fissure.
8		Sealed behind a Dendro rock in Debris of Panjvahe. Use Dendrograma to break it.
9		Near an air draft, east of the waypoint in Debris of Panjvahe. Use the air draft to reach higher ground.
10		Floating across the Shrine of Depth in Debris of Panjvahe. Glide across from a higher point.
11		By a high cliff just above the waypoint north of The Sands of Three Canals. Climb the cliff.
12		On top of a cactus northwest of The Sands of Three Canals. Climb the cactus.
13		In the same area as the Primal Construct battle arena of Dune-Emtombed Fecundity: Part 2.
14		High above The Sands of Three Canals. Activate the Anemo Monument for updraft. Use Four-Leaf Sigil to the east to reach an invisible platform.
15		Sealed behind a Dendro rock in The Sands of Three Canals. Use Dendrograna to break it.
16		Below the waypoint west of The Sands of Three Canals, hanging by a branch. Glide down or climb from below.
17		On top of a dead tree at The Sands of Three Canals. Climb the dead tree.
18		Inside a tent south of The Sands of Three Canals. Enter the tent.
19		Sealed inside a Dendro rock in The Sands of Three Canals. Use Liloupar gadget to unlock it. Dendrograma branch is nearby.
20		Floating high above the valley west of The Sands of Three Canals. Use the cliff by the huge branch. An invisible bridge will appear.
21		On top of a dead tree at Wounded Shin Valley. Climb the dead tree.
22		Inside the hollow head of the Ruin Golem. Dune-Entombed Fecundity: Part 3 quest of The Dirge of Bilqis.
23		In the area where you obtain Liloupar’s fragment during Dune-Entombed Fecundity: Part 3 quest of The Dirge of Bilqis.
24		Deep inside a small cave in the Ruin Golem tunnels. Dune-Entombed Fecundity: Part 3 quest of The Dirge of Bilqis.
25		On top of a dead tree between Wounded Shin Valley and The Temples Forsaken. Climb the dead tree.
26		Inside the small temple northwest of The Temples Forsaken, floating by the Four-Leaf Sigil. Use the sigil.
27		Next to the waypoint west of Safhe Shatranj by the Four-Leaf Sigil. Use the sigil.
28		Obtainable after sand recedes for completing ruins in The Temple Where Sand Flows Like Tears quest of The Dirge of Bilqis.
29		Outside The Temples Forsaken, near the east waypoint. Explore the area around the waypoint.
30		Inside a small cave outside The Temple Forsaken. Enter the small cave.
31		Inside a small cave right above the Four-Leaf Sigil near The Temples Forsaken. Use the sigil to reach the cave.
32		On top of a dead tree at the Tanit Camps. Climb the dead tree.
33		On top of a broken pillar beneath the waypoint east of Wadi Al-Majuj. Climb the broken pillar.
34		Underground area northwest of Wadi Al-Majuj, near the top of a large statue. Explore the underground area.
35		Inside the ruins after completing the Apocalypse Lost quest. Go up an invisible ramp inside.
36		Hanging on a branch north of Wadi Al-Majuj. Glide from a higher point to the branch.
37		On top of a broken pillar beneath the waypoint east of Wadi Al-Majuj. Climb the broken pillar.
38		Near the center of the Eternal Oasis. Locked behind The Eternal Dream, Ever Lush quest of The Dirge of Bilqis.
39		By the grassy area of the river shore in Eternal Oasis. Locked behind The Eternal Dream, Ever Lush quest of The Dirge of Bilqis.
40		Inside an Atmospheric Vortex in Eternal Oasis. Use max level Liloupar gadget upgraded from Tadhla’s quest chain.
41		Atop a tree, above an Eremite camp in Passage of Ghouls. Climb the tree.
42		Floating on a large rock, down the fissure in Passage of Ghouls. Glide down into the fissure.
43		On top of the tree, south of the Qusayr Al-Inkhida’ waypoint. Climb the tree.
44		Use Four-Leaf Sigil to reach the top of the room in Qusayr Al-Inkhida’. Waypoint locked behind Dune-Entombed Fecundity: Part 1 quest.
45		High in the air near Qusayr Al-Inkhida’. Use Dendrograma to break Dendro rocks and reveal Dendro monuments. Light all four for a wind current.
46		Hidden inside a tree trunk inside the Eternal Oasis. Locked behind The Eternal Dream, Ever Lush quest of The Dirge of Bilqis.
47		On top of a cactus, south of The Sands of Three Canals. Climb the cactus.
48		Sealed behind a Dendro rock. Use Dendrograma to break it.
49		Floating above the Setekh Wenut tunnel entrance. Glide down from a higher point.
50		Underground near the City of Deceased domain, hidden behind a pile of rocks. Break the rocks.
51		Investigate a rock formation inside the Wenut Tunnels to reveal a hole. Look into the hole from the northern ledge.
52		Follow The Temple Where Sand Flows Like Tears quest of The Dirge of Bilqis to the second Ruin Mechanism room. Atop a large branch.
53		Blow off the sand pile on the mechanism and activate it to reveal a hole. During The Temple Where Sand Flows Like Tears quest.
54		Begin The Temple Where Sand Flows Like Tears quest to enter the temple. Solve Rune Mechanism puzzles to see it.
55		Inside the cavern, atop branches inside a ruin door, during Dune-Entombed Fecundy: Part 2 quest.

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Girdle of the Sands Dendroculus Locations {#girdle-of-the-sands-dendroculus-locations}

No.	Image	How to Get
1		On top of a hut in a Hilichurl camp west of the waypoint. Climb the hut.
2		On a cactus above the Molten Iron Fortress Domain. Climb the cactus.
3		At the tip of a pole next to an Anemo Hilichurl Rogue. Defeat the enemy or avoid it.
4		In an underground area at Temir Mountains. Near the bottom of the shaft. Explore the underground area.
5		At the tip of a gray crystal in Temir Mountains. Climb the crystal.
6		On top of a large forge found during the Khvarena of Good and Evil quest.
7		Along the edge of the cliff. Use Four-Leaf Sigils to gain elevation, then climb the cliff.
8		At the tip of a large gray crystal surrounding Tunigi Hollow. Climb the crystal.
9		Underground by a cliff with Four-Leaf Sigils. Requires Khvarena of Good and Evil quest progression.
10		At the center of Tunigi Hollow.
11		Under the elevator after connecting three Mysterious Energy Receivers during Khvarena of Good and Evil quest.
12		On a large gray crystal east of Tunigi Hollow. Get on top of the east cliff and glide down.
13		Near the ceiling inside a room with wind currents and sunlight. Khvarena of Good and Evil quest progression required.
14		Under an arch just below a Four-Leaf Sigil. Use the sigil and look under the arch.
15		Above a tree high up a cliff with a Consecrated Red Vulture nearby. Defeat or avoid the enemy.
16		Inside a tree stump. Enter the tree stump.
17		Floating in the air near a high cliff. Use Sorush to adjust the Four-Leaf Sigil.
18		On top of a giant tree root. Climb the tree root.
19		Floating in the air between branches. Climb to the highest point on the tree and glide.
20		Floating in the air. Climb the nearby cliff and glide down.
21		Floating on top of a rock. Get to higher ground and glide down.
22		Floating beneath a small rock bridge, in an opening leading underground.
23		On top of a small tree next to a Soul Bell. Climb the small tree.
24		Sitting on top of a Treasure Hoarder hut by the cliff. Climb the hut.
25		On top of a large tree. Climb the large tree.
26		On top of a large branch. Climb onto the large branch.
27		Above the water. Glide from one of the cliffs in the Samudra Coast.
28		Inside an underground arena found during Khvarena of Good and Evil quest.
29		Underground in an area with Mysterious Energy Control Systems. Take the elevator up to an area with Ruin Guards.
30		Behind a gray crystal entrance inside an underground cave. Break the crystal or find the entrance.
31		Between the top of a large gray crystal and a cliff. Climb the crystal and cliff.
32		Beneath the edge of a large cliff. Look over the edge of the cliff.
33		On top of a large spike in the underground cave at the Gate of Zulqarnain. Follow the road north.
34		Beneath a large cliff with a Consecrated Red Vulture on top. Defeat or avoid the enemy.
35		On top of a hut in a Hilichurl camp west of the Gate of Zulqarnain waypoint. Climb the hut.
36		Inside a broken piece of machinery next to Ruin Drakes. Defeat or avoid the enemies.

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How to Use the Dendroculus

Upgrade the Sumeru Statue of the Seven

Dendroculi are collectible items in Sumeru used to level up the Statues of the Seven. Upgrading these statues provides rewards such as Primogems, Shrine of Depths Keys, and Adventure EXP, which are essential for game progression.

Extra Dendroculus

It’s common to have 1 extra Dendroculus remaining after maxing out the Statues. Currently, there are no known uses for this extra Dendroculus, but it serves as confirmation you’ve collected all available in the current version.

Learn How to Level Up Statues of the Seven

Explore Sumeru with Friends in Co-op

Exploring Sumeru to find all Dendroculus can be challenging. Consider inviting friends to join your world in co-op mode! Team up to explore, solve puzzles, and locate these collectibles more efficiently. Find fellow travelers on our friend request and co-op boards:

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Supporting Cast Board

Rewards from Sumeru Statue of the Seven Upgrades

Lv.	Dendroculus Required	Rewards
2	20	Adventure EXP x80 Primogem x60 Dendro Sigil x5 Sumeru Shrine of Depths Key x1
3	22	Adventure EXP x120 Primogem x60 Dendro Sigil x5 Memory of Flourishing Green x1
4	25	Adventure EXP x160 Primogem x60 Dendro Sigil x5 Sumeru Shrine of Depths Key x1
5	28	Adventure EXP x200 Primogem x60 Dendro Sigil x10 Memory of Flourishing Green x1
6	30	Adventure EXP x240 Primogem x60 Dendro Sigil x10 Sumeru Shrine of Depths Key x1
7	32	Adventure EXP x280 Primogem x60 Dendro Sigil x10 Memory of Flourishing Green x1
8	35	Adventure EXP x320 Primogem x60 Dendro Sigil x15 Sumeru Shrine of Depths Key x1
9	38	Adventure EXP x360 Primogem x60 Dendro Sigil x15 Sumeru Shrine of Depths Key x1
10	40	Adventure EXP x400 Primogem x60 Dendro Sigil x15 Sumeru Shrine of Depths Key x1

Dendroculus Resonance Stone Gadget

Locate Missing Dendroculus with Resonance Stones

If you’re struggling to find the last few Dendroculus, the Dendroculus Resonance Stone is an invaluable tool. This gadget helps pinpoint the general area of any remaining Dendroculus in your vicinity. The blueprint for crafting the Dendroculus Resonance Stone is obtainable as a reward from increasing your City Reputation in Sumeru.

Learn How to Craft and Use the Dendroculus Resonance Stone

Using the Dendroculus Resonance Stone

Once crafted, equip the Dendroculus Resonance Stone from your inventory. Upon use, it will display a highlighted area on your map, indicating the approximate location of an uncollected Dendroculus. Explore this marked zone to find your missing collectible.

Detailed Oculus Resonance Stone Guide

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